There’s something strange about Freud’s _Civilization and its Discontents_ (1930). Biologically, Freud was a Neo-Lamarckian, who believed in both the modification of organisms through need and the inheritance of acquired characteristics. However, in _Civilization_, Freud argued that because human nature is immutable, society has dim odds of improving substantially. Lamarckians, of course, rejected that any species-nature is immutable, as species can always be transformed via the inheritance of acquired characteristics. In fact, many of Freud’s Viennese contemporaries—such as Wilhelm Reich, Julius (...) Tandler, and Paul Kammerer—took their Lamarckism to license precisely the sorts of radical social projects Freud deemed impossible. Thus the Freud of _Civilization_ helped himself to a rigid view of human nature which, given his associated biological views, he seemingly ought to have rejected. In this paper, I explain this apparent inconsistency, and suggest Freud resolved it in the following way: Freud was not merely a Lamarckian, but also a strong and peculiar kind of recapitulationist, who believed stages of psychological development both recapitulate phylogeny, and “remain with us” throughout both individual lives and future species-history. I suggest Freud’s recapitulationism supposed a certain inertia: what occurred in phylogenetic history cannot un-occur, and therefore there are aspects of our nature which we cannot un-acquire. In this way, Freud reached a rigid conception of human nature despite his Lamarckism. (shrink)

Electronics can be defined as electromagnetic technology dealing with information, and meta-electronics as a field encompassing all the synergistic technologies in which electronics plays a dominant role. Examining the broad field corresponding to this definition we realize that its history starts some seventy years earlier than the customarily accepted birth of electronics, and, what is more significant, that electronics undergoes a true evolution. This new evolution creates rich, diverse structures similar to those created by the biological evolution. Like biology, electronics (...) is non-teleological, which allows for its unlimited evolutionary development. We propose electronic analogies of all essential biological categories, at all levels: population, speciation, common ancestor, phenotype, extended phenotype, co-evolution, convergent evolution, evolutionary arms race, extinction and mass extinction, hierarchical levels, generative entrenchment, genes, alleles, genome, genetic pool, recombination, mutation, genetic drift, lateral gene transfer, etc. The evolutionary algorithm operating in electronics, like a Darwinian one, includes variation within a population of device models, heredity, natural selection, and a form of selection based on aesthetics and fashion which resembles sexual selection. Algorithm is especially similar to artificial selection, thus possessing directionality in the variational part. Electronic development is orders of magnitude faster than biological, accelerated by that directionality and by other distinct, identifiable mechanisms. Speciation in electronics, as in biology, is best represented on a phylogenetic tree, which starts from a common ancestor, but lately exhibits a unification trend. If continued, this trend may lead to the appearance of a common descendant absent in biology. Our analysis may explain emerging anti-social aspects of electronics and our conclusions add new urgency to recent concerns with unchecked development of Artificial Intelligence. (shrink)

The bipolar type systems in the sense ofKretschmer, Jaensch, Jung and psychology of form current in presentday psychology are subjected to criticism from the point of view of their practical utility for the diagnosis of human beings, since they are unable to do justice to the complexity of human nature. The author sees two chief problems in the concept of type. According to their function, types are comparable with ultimate forms in which conclusions are drawn from a number of familiar (...) characteristics as to the simultaneous occurrence of the remaining characteristics which are more difficult to apprehend empirically. Their nature places types in close analogy to forms and melodies. Assignment to certain types is not a mere classification, but a heuristic speculation. The response of primitive organisms to complexes of characteristics represents a phylogenetically older form of typological conclusion. Special emphasis is laid upon the intimate relation existing between the typological method of apprehension and the active side of organisms. Brief reference is made to the way in which types can be statistically derived, the apriority problem of the type and the transmission of types by heredity.Les systèmes de types bipolaires usuels dans la psychologie actuelle d'aprèsKretschmer, Jaensch etJung et la psychologie de la structure de formes sont soumis à une critique établie du point de vue de son application pratique au diagnostic humain, car ils ne permettent pas de venir à bout de la nature humaine. L'auteur voit dans la notion des types deux problèmes principaux: d'après leur fonction les types sont comparables à des syllogismes dans lesquelles à l'intérieur d'une série de caractères associés on déduit d'un certain nombre de caractères connus la présence de caractères individuels difficiles à saisir empiriquement. D'après leur nature les types sont en analogie avec les figures et les mélodies. L'ordonnancement des types n'est pas une simple classification mais une entreprise heuristique. La recherche des complexes de caractères dans les organismes primitifs représente une forme phylogénétiquement ancienne de la déduction typologique. La relation étroite entre le mode de conception typologique et la manière d'agir des organismes est particulièrement souligné. La déductibilité statistique, la question d'apriorité des types et leur hérédité sont brièvement effleurées. (shrink)

The dichotomy between Nature and Nurture, which has been dismantled within the framework of development, remains embodied in the notions of plasticity and evolvability. We argue that plasticity and evolvability, like development and heredity, are neither dichotomous nor distinct: the very same mechanisms may be involved in both, and the research perspective chosen depends to a large extent on the type of problem being explored and the kinds of questions being asked. Epigenetic inheritance leads to transgenerationally extended plasticity, and (...) developmentally-induced heritable epigenetic variations provide additional foci for selection that can lead to evolutionary change. Moreover, hereditary innovations may result from developmentally induced large-scale genomic repatterning events, which are akin to Goldschmidtian “systemic mutations”. The epigenetic mechanisms involved in repatterning can be activated by both environmental and genomic stress, and lead to phylogenetic as well as ontogenetic changes. Hence, the effects and the mechanisms of plasticity directly contribute to evolvability. (shrink)

This article re-contextualizes Sigmund Freud's interest in the idea of the inheritance of acquired characteristics in terms of the socio-political connotations of Lamarckism and Darwinism in the 1930s and 1950s. Many scholars have speculated as to why Freud continued to insist on a supposedly outmoded theory of evolution in the 1930s even as he was aware that it was no longer tenable. While Freud's initial interest in the inheritance of phylogenetic memory was not necessarily politically motivated, his refusal to (...) abandon this theory in the 1930s must be understood in terms of wider debates, especially regarding the position of the Jewish people in Germany and Austria. Freud became uneasy about the inheritance of memory not because it was scientifically disproven, but because it had become politically charged and suspiciously regarded by the Nazis as Bolshevik and Jewish. Where Freud seemed to use the idea of inherited memory as a way of universalizing his theory beyond the individual cultural milieu of his mostly Jewish patients, such a notion of universal science itself became politically charged and identified as particularly Jewish. The vexed and speculative interpretations of Freud's Lamarckism are situated as part of a larger post-War cultural reaction against Communism on the one hand, and on the other hand, against any scientific concepts of race in the wake of World War II. (shrink)

Feinberg and Mallatt, in their presentation of neurobiological naturalism, have suggested that visual consciousness was acquired by early vertebrates and inherited by a wide range of descendants, and that its neural basis has shifted to nonhomologous nervous structures during evolution. However, their evolutionary scenario of visual consciousness relies on the assumption that visual consciousness is closely linked with survival, which is not commonly accepted in current consciousness research. We suggest an alternative idea that visual consciousness is linked to a specific (...) class of agency, consequently justifying their phylogenetic claim. We also examine the implication of their phylogenetic claim: visual consciousness is homologous across vertebrates, but its neural basis is not. This apparent incongruence illustrates a general phenomenon of homology, and that the resulting hierarchical view of visual consciousness and its neural basis can be straightforwardly accommodated by neurobiological naturalism. Throughout these discussions, we aim to address the potential theoretical issues in neurobiological naturalism and refine the picture illustrated by Feinberg and Mallatt regarding phylogenetic distribution and trajectories of visual consciousness. (shrink)

Heredity has been dismissed as an insignificant object in Claude Bernard’s physiology, and the topic is usually ignored by historians. Yet, thirty years ago, Jean Gayon demonstrated that Bernard did elaborate on the subject. The present paper aims at reassessing the issue of heredity in Claude Bernard’s project of a “general physiology”. My first claim is that Bernard’s interest in heredity was linked to his ambitious goal of redefining general physiology in relation to morphology. In 1867, not (...) only was morphology included within experimental physiology, but it also theoretically grounded physiological investigations. By 1878, morphology and physiology were considered as completely independent sciences, and only the latter was perceived as suitable to experimentation. My second claim is that this reversal reflected the existence of two opposite attitudes towards heredity. In the late 1860s, Bernard was convinced that heredity would soon be accessible to experimental manipulation and that new species would be produced in the laboratory exactly like organic chemistry succeeded to do for raw bodies. Ten years later, he ascertained that this was impossible. My third claim is that Bernard was epistemologically ill-equipped to address the issue of heredity. Bernard was strongly committed to a general reasoning scheme that acknowledged only three categories: determining conditions, constant laws and phenomena. This scheme was a key factor in his successes as a physiologist who was able to capture new mechanisms in living bodies. Nonetheless, it also prevented him from understanding how time and history could be endowed with a causal action that cannot be reduced to timeless parameters. (shrink)

The long-awaited revision of the industry standard on phylogenetics Since the publication of the first edition of this landmark volume more than twenty-five years ago, phylogenetic systematics has taken its place as the dominant paradigm of systematic biology. It has profoundly influenced the way scientists study evolution, and has seen many theoretical and technical advances as the field has continued to grow. It goes almost without saying that the next twenty-five years of phylogenetic research will prove as fascinating (...) as the first, with many exciting developments yet to come. This new edition of Phylogenetics captures the very essence of this rapidly evolving discipline. Written for the practicing systematist and phylogeneticist, it addresses both the philosophical and technical issues of the field, as well as surveys general practices in taxonomy. Major sections of the book deal with the nature of species and higher taxa, homology and characters, trees and tree graphs, and biogeography—the purpose being to develop biologically relevant species, character, tree, and biogeographic concepts that can be applied fruitfully to phylogenetics. The book then turns its focus to phylogenetic trees, including an in-depth guide to tree-building algorithms. Additional coverage includes: Parsimony and parsimony analysis Parametric phylogenetics including maximum likelihood and Bayesian approaches Phylogenetic classification Critiques of evolutionary taxonomy, phenetics, and transformed cladistics Specimen selection, field collecting, and curating Systematic publication and the rules of nomenclature Providing a thorough synthesis of the field, this important update to Phylogenetics is essential for students and researchers in the areas of evolutionary biology, molecular evolution, genetics and evolutionary genetics, paleontology, physical anthropology, and zoology. (shrink)

This commentary acknowledges the importance of Davey's biocognitive approach to the uneven distribution of fears on the basis of its contribution to a human model for understanding fear. An integrated heredity-environment and developmental transactional approach based on field/system theory is recommended in place of the mechanistic heredity × environment interactionism that Davey uses to explain behavioral ontogeny.

Argues for the primacy of the phylogenetic system as the general reference system in biology. This book, first published in 1966, generated significant controversy and opened possibilities for evolutionary biology.

Excerpt from Heredity, Correlation and Sex Differences in School Abilities: Studies From the Department of Educational Psychology at Teachers College, Columbia University The Relationships between the Different Abilities Involved in the Study of Arithmetic. By W. A. Fox and E. L. Thorndike. About the Publisher Forgotten Books publishes hundreds of thousands of rare and classic books. Find more at www.forgottenbooks.com This book is a reproduction of an important historical work. Forgotten Books uses state-of-the-art technology to digitally reconstruct the work, (...) preserving the original format whilst repairing imperfections present in the aged copy. In rare cases, an imperfection in the original, such as a blemish or missing page, may be replicated in our edition. We do, however, repair the vast majority of imperfections successfully; any imperfections that remain are intentionally left to preserve the state of such historical works. (shrink)

Phylogenetics has inherent utility in evolutionary developmental biology (EDB) as it is an established methodology for estimating evolutionary relationships and for making comparisons between levels of biological organization. However, explicit phylogenetic methods generally have been limited to two levels of organization in EDB—the species and the gene. We demonstrate that phylogenetic methods can be applied broadly to other organizational levels, such as morphological structures or cell types, to identify evolutionary patterns. We present examples at and between different hierarchical (...) levels of organization to address questions central to EDB. We argue that this application of “hierarchical phylogenetics” can be a unifying analytical approach to the field of EDB. BioEssays 27:1158–1166, 2005. © 2005 Wiley Periodicals, Inc. (shrink)

Cadherins are a multigene family of proteins which mediate homophilic calcium‐dependent cell adhesion and are thought to play an important role in morphogenesis by mediating specific intercellular adhesion. Different lines of experimental evidence have recently indicated that the site responsible for mediating adhesive interactions is localized to the first extracellular domain of cadherin. Based upon an analysis of the sequence of this domain, I show that cadherins can be classified into three groups with distinct structural features. Furthermore, using this sequence (...) information a phylogenetic tree relating the known cadherins was assembled. This is the first such tree to be published for the cadherins. One cadherin subtype, neural cadherin (N‐cadherin), shows very little sequence divergence between species, whereas all other cadherin subtypes show more substantial divergence, suggesting that selective pressure upon this domain may be greater for N‐cadherin than for other cadherins. Phylogenetic analysis also suggests that the gene duplications which established the main branches leading to the different cadherin subtypes occurred very early in their history. These duplications set the stage for the diversified superfamily we now observe. (shrink)

This paper aims to give an impression of how biologists, at the turn of the twentieth century, came to conceptualize and define the hidden entities presumed to govern the process of hereditary transmission. With that, the stage was set for the emergence of genetics as a biological discipline that came to dominate the life sciences of the twentieth century. The annus mirabilis of 1900, with its triple re-appreciation of Gregor Mendel’s work by the botanists Hugo de Vries, Carl Correns, and (...) Erich Tschermak, can be seen as the watershed after which theorizing about heredity and experimentation—selecting pure lines and Mendelian crossing—became tightly connected. As to concepts before this, this paper will analyze Carl von Nägeli’s ‘idioplasm’, Hugo de Vries’s ‘pangenes’, and August Weismann’s ‘germ plasm’. Carl Correns’s Anlagen and Wilhelm Johannsen’s ‘genes’ would replace them in the decade after 1900.Keywords: Idioplasm ; Pangenes ; Germ plasm ; Anlagen ; Gene; Genotype. (shrink)

The concept of ‘phylogenetic inertia’ is routinely deployed in evolutionary biology as an alternative to natural selection for explaining the persistence of characteristics that appear sub-optimal from an adaptationist perspective. However, in many of these contexts the precise meaning of ‘phylogenetic inertia’ and its relationship to selection are far from clear. After tracing the history of the concept of ‘inertia’ in evolutionary biology, I argue that treating phylogenetic inertia and natural selection as alternative explanations is mistaken because (...) phylogenetic inertia is, from a Darwinian point of view, simply an expected effect of selection. Although Darwin did not discuss ‘phylogenetic inertia,’ he did assert the explanatory priority of selection over descent. An analysis of ‘phylogenetic inertia’ provides a perspective from which to assess Darwin’s view. (shrink)

The aptationist program includes attempts at sorting adaptations from exaptations, and therefore requires knowledge of historical changes in biological character states (traits) and their effects or functions, particularly for nonoptimal aptations. Phylogenetic inference is a key approach for historical aspects of evolutionary hypotheses, particularly testing evolutionary scenarios, and such “tree-thinking” investigation is directly relevant to the aptationist program.

This chapter develops a subtle model that integrates environmental and internal factors. It describes the phylogenetic distribution of multicellular organisms in general and complex multicellular life in particular, clarifying the important distinction between the two. This chapter shows that the long apparent lag between the appearance of simple multicellularity in eukaryotes and the radiation of groups with complex multicellular organization has an environmental component that can be associated back to the consequences of life with interior and exterior cells. It (...) suggests that the evolutionary transition from unicells to complex multicellular organisms has several steps. (shrink)

define the hidden entities presumed to govern the process of hereditary transmissionWith that Hans-Jörg came to conceptualize, Carl Correns, its triple re-appreciation of Gregor Mendel’s work by the botanists Hugo de Vries, Erich Tschermak can be seen as the watershed after which theorizing about heredity & Pure Experimentation—Selecting.

The occurrence of hiccoughs (hiccups) is very widespread and yet their neuronal origin and physiological significance are still unresolved. Several hypotheses have been proposed. Here we consider a phylogenetic perspective, starting from the concept that the ventilatory central pattern generator of lower vertebrates provides the base upon which central pattern generators of higher vertebrates develop. Hiccoughs are characterized by glottal closure during inspiration and by early development in relation to lung ventilation. They are inhibited when the concentration of inhaled (...) CO2 is increased and they can be abolished by the drug baclofen (an agonist of the GABAB receptor). These properties are shared by ventilatory motor patterns of lower vertebrates, leading to the hypothesis that hiccough is the expression of archaic motor patterns and particularly the motor pattern of gill ventilation in bimodal breathers such as most frogs. A circuit that can generate hiccoughs may persist in mammals because it has permitted the development of pattern generators for other useful functions of the pharynx and chest wall muscles, such as suckling or eupneic breathing. BioEssays 25:182–188, 2003. © 2003 Wiley Periodicals, Inc. (shrink)

Historians have emphasized the appearance of a productive research program in genetics after 1910, and philosophers and biologists have considered endorsement of genetics as a progressive move, indeed as a starting point for modern experimental biology. These efforts focus on what biology had changed to. This paper examines the condition from which biology moved, stressing the way in which Americans held heredity and development as a natural, intimately intertwined couple. Heredity accounts for likenesses, development for variation, and the (...) two act together throughout an individual's life. This discussion establishes the antecedent conditions necessary to explain the split of genetics and embryology and addresses the reasons for the split. (shrink)

This study explores the principles of meritocracy and heredity as formulated in the three works of early Daoist philosophy, the Laozi, Zhuangzi and Wenzi. Because Daoist philosophy emerged in critical response to the Confucian worldview, this investigation is placed against the backdrop of pertinent Confucian propositions. To this end, the study begins with a review of Confucian positions on the issue of meritocracy and heredity as expressed in the main transmitted works, as well as newly excavated texts that (...) can be associated with this intellectual tradition. The paper concludes that, while indeed rejecting the Confucian understanding of meritocracy, Daoist texts operated with their own concepts on human excellency. Moreover, the opposition between meritocracy and heredity characteristic of some Confucian works appears less pronounced in early Daoist philosophy. (shrink)

This paper argues that our modern concept of biological heredity was first clearly introduced in a theoretical and practical setting by the generation of French physicians that were active between 1810 and 1830. It describes how from a traditional focus on hereditary transmission of disease, influential French medical men like Esquirol, Fodéré, Piorry, Lévy, moved towards considering heredity a central concept for the conception of the human bodily frame, and its set of physical and moral dispositions. The notion (...) of heredity as a natural force, with a wide ranging capabilities of transmitting differentially both fundamental and accidental characters was generalized by that generation of physicians with the help of contemporary naturalists and physiologists. By 1830 the term hérédité was widespread, and it shared the explanatory and semantic qualities of traditional medical concepts like constitution and temperament. An analysis is given of the main developments that led to the conception of biological (including human) bodies as consisting of a layered, hierarchical organization of characters, differentially affected by the laws of conservation (Heredity) and change (Inneity, Variation). The mid-century work of the French physician Prosper Lucas, Traité Philosophique et Physiologique de L'Hérédité Naturelle, is shown to be the culmination of the efforts of several generations of French physicians towards having a feasible, complexly structured notion of how heredity works. (shrink)

This book, first published in 1938, is based on the Muirhead Lectures given at Birmingham University in February and March of 1937. The first half of this book is mainly devoted to an exposition of the principles of genetics, whilst the second half deals with more controversial topics, with the text providing an insight into the ideology of the time. This title will be of interest to students of politics and history.

The concept of a temporal phylogenetic network is a mathematical model of evolution of a family of natural languages. It takes into account the fact that languages can trade their characteristics with each other when linguistic communities are in contact, and also that a contact is only possible when the languages are spoken at the same time. We show how computational methods of answer set programming and constraint logic programming can be used to generate plausible conjectures about contacts between (...) prehistoric linguistic communities, and illustrate our approach by applying it to the evolutionary history of Indo-European languages. (shrink)

The author proposes a system of identification, positional, and phyletic numbers for taxa that makes possible a significant relationship between numerical classification and phylogeny.

This review paper discusses the perspective of complex biological systems as applied to inheritance and ontogeny, focusing on the continuity of genetic, epigenetic and microbiotic inheritance. The informational processuality within this continuity can be used as to exemplify the insufficiency of hierarchical concepts in grasping the complex and integrated nature of biological processes. The argument follows Bruni and Giorgi in emphasizing that while structures and substrates are organized hierarchically, communicational processes are organized heterarchically. The essay also argues the insufficiency of (...) a single, basic, i.e. genetic level of description, which is the prevalent idea of twentieth century biology, to explain all phenotypic variation. I argue that inheritance and development cannot be fully explained by some sub- or super-ordination and that such descriptions are merely heuristic tools that do not reflect the nature of such processes. (shrink)

The Norwegian Association for Heredity Research played a key role in the rise of genetics as a research field in Norway. The immediate background of its establishment in 1919 was the need for an organization that could clarify scientific issues regarding eugenics and coordinate Norwegian representation in the organized international eugenics movement. The Association never assumed this role. Instead, Norway was represented in the international eugenics movement by the so-called Norwegian Consultative Eugenics Commission, whose leader, Jon Alfred Mjøen, was (...) dismissed as a pseudo-scientist by Norwegian geneticists. The paper explores the Association’s role in defining and delimiting scientific expert knowledge in the field of genetics and eugenics in Norway. It demonstrates how struggles about academic authority on the national arena were intertwined with struggles about representation and impact in the international eugenics movement and how transnational scientific networks where mobilized to legitimize and delegitimize notions about Nordic race supremacy, racial mixing and the politics of eugenic sterilizations. (shrink)

Darwin's ideas on variation, heredity, and development differ significantly from twentieth-century views. First, Darwin held that environmental changes, acting either on the reproductive organs or the body, were necessary to generate variation. Second, heredity was a developmental, not a transmissional, process; variation was a change in the developmental process of change. An analysis of Darwin's elaboration and modification of these two positions from his early notebooks (1836-1844) to the last edition of the /Variation of Animals and Plants Under (...) Domestication/ (1875) complements previous Darwin scholarship on these issues. Included in this analysis is a description of the way Darwin employed the distinction between transmission and development, as well as the conceptual relationship he saw between heredity and variation. This paper is part of a larger project comparing commitments regarding variation during the latter half of the nineteenth century. (shrink)

Did Man Get Here by Evolution or by Creation? is a book whose publication date, 1967, indicates that the doctrine of evolution is still being rejected today by at least some segments of the intellectual community. As reverberations of the nationally famous Scopes trial continue, it may be profitable to inquire into the principles which contemporary biologists utilize in explaining the general course of evolution. For it would seem that, if these scientific principles were sufficiently cogent, there could be no (...) valid reason for questioning the doctrine. (shrink)

Amine acetylation is a diverse topic with importance to the regulation of several physiological processes as well as the metabolism of drugs and environmental chemicals. Arylalkylamine N‐acetyltransferase is widely distributed in several species, where this enzyme plays an important role in the seasonal regulation of reproduction and photoperiodism in vertebrates through the pathway of melatonin formation. In insects, this enzyme is involved in monoamine neurotransmitter inactivation and the formation of catecholamine intermediates necessary for sclerotization of the insect cuticle.

This chapter contains section titled: The Relation of Genotype to Phenotype Statistical Approaches to the Study of Quantitative Characters Problems Raised by Statistical Methodologies Making Quantitative Trait Genes Real Bibliography.

On the classical account of evolution by natural selection found in Lewontin and many subsequent authors, ENS is conceived as involving three key ingredients: phenotypic variation, fitness differences, and heredity. Through the analysis of three problem cases involving heredity, I argue that the classical conception is substantially flawed, showing that heredity is not required for selection. I consider further problems with the classical account of ENS arising from conflations between three distinct senses of the central concept of (...) ‘fitness’ and offer an alternative to the classical conception of ENS involving the interaction of distinct evolutionary mechanisms. (shrink)

We develop a conceptual framework that connects biological heredity and organization. We refer to heredity as the cross-generation conservation of functional elements, defined as constraints subject to organizational closure. While hereditary objects are functional constituents of biological systems, any other entity that is stable across generations—and possibly involved in the recurrence of phenotypes—belongs to their environment. The central outcome of the organizational perspective consists in extending the scope of heredity beyond the genetic domain without merging it with (...) the broad category of cross-generation stability. After discussing some implications, we conclude with a reflection on the relationship between stability and variation. 1Introduction2From Extended Heredity to Cross-generation Stability 2.1Extending the scope of heredity: A brief state of the art2.2Rethinking heredity: Conceptual challenges3Biological Heredity in Light of Organization 3.1Biological organization within organisms and beyond3.2Extending organization in time3.3What is biological heredity?4Implications and Objections 4.1Heredity as a specific kind of cross-generation stability4.2Heredity at various levels of description4.3Non-functional and dysfunctional objects5Conclusions: From Conservation to Variation. (shrink)