Abstract
This article focuses on two indices from Geschlecht III session XIII: (1) an apparently insignificant reference to Stiegler and (2) the recourse to the concept of the engram as a trope of other grammatological figures that are more frequent in Derrida's work. By interweaving these indexes together, the article suggests that Derrida's text can be read as a noteworthy stage in his ongoing dialogue with Bernard Stiegler surrounding the question posed by human evolution to any accounts of the history of life. Along this path, the article inscribes Derrida's (en-)grammatological history of life within the line of thought that goes from Richard Semon's engram theory to Simona Ginsburg and Eva Jablonka's contemporary re-elaboration of this theory. In doing so, it argues that, although for Stiegler Derrida's grammatology aligns with biological reductionism, the latter may provide the theoretical framework for current evolutionary accounts of life as plasticity.