Contemporary evolutionary biology comprises a plural landscape of multiple co-existent conceptual frameworks and strenuous voices that disagree on the nature and scope of evolutionary theory. Since the mid-eighties, some of these conceptual frameworks have denounced the ontologies of the Modern Synthesis and of the updated Standard Theory of Evolution as unfinished or even flawed. In this paper, we analyze and compare two of those conceptual frameworks, namely Niles Eldredge’s Hierarchy Theory of Evolution (with its extended ontology of evolutionary (...) entities) and the Extended Evolutionary Synthesis (with its proposal of an extended ontology of evolutionary processes), in an attempt to map some epistemic bridges (e.g. compatible views of causation; niche construction) and some conceptual rifts (e.g. extra-genetic inheritance; different perspectives on macroevolution; contrasting standpoints held in the “externalism–internalism” debate) that exist between them. This paper seeks to encourage theoretical, philosophical and historiographical discussions about pluralism or the possible unification of contemporary evolutionary biology. (shrink)

This paper examines the subversive role of statistics paleontology at the end of the 19th and the beginning of the 20th centuries. In particular, I will focus on German paleontology and its relationship with statistics. I argue that in paleontology, the quantitative method was questioned and strongly limited by the first decade of the 20th century because, as its opponents noted, when the fossil record is treated statistically, it was found to generate results openly in conflict with the Darwinian theory (...) of evolution. Essentially, statistics questions the gradual mode of evolution and the role of natural selection. The main objections to statistics were addressed during the meetings at the Kaiserlich-Königliche Geologische Reichsanstalt in Vienna in the 1880s. After having introduced the statistical treatment of the fossil record, I will use the works of Charles Léo Lesquereux, Joachim Barrande, and Henry Shaler Williams to compare the objections raised in Vienna with how the statistical treatment of the data worked in practice. Furthermore, I will discuss the criticisms of Melchior Neumayr, one of the leading German opponents of statistical paleontology, to show why, and to what extent, statistics were questioned in Vienna. The final part of this paper considers what paleontologists can derive from a statistical notion of data: the necessity of opening a discussion about the completeness and nature of the paleontological data. The Vienna discussion about which method paleontologists should follow offers an interesting case study in order to understand the epistemic tensions within paleontology surrounding Darwin’s theory as well as the variety of non-Darwinian alternatives that emerged from the statistical treatment of the fossil record at the end of the 19th century. (shrink)

Dr. Marjorie Grene has argued that criteria taken from a personalist philosophy of science have regulative force in the dispute between orthogenetic and synthetic or neo-Darwinian theories of evolution, and that these criteria commend the acceptance of the orthogenetic position. Grene's position includes two basically correct theses concerning the limitations of operationism and reductionism. However, she fails to show that personalist tenets are necessary for the validation of these two theses. Moreover, the proposed modifications of evolutionary theory depend upon (...) additional premisses: that biology must study individuals rather than populations, and that the synthetic theory must prove that natural selection and mutation are the only possible factors for control of the direction of evolutionary change. The evidence for these premisses is called into question. (shrink)

After the theory of evolution was put forward by Charles Darwin, it has been interpreted in terms of different philosophical paradigms. The most important of these interpretations stand out as the intelligent designers and the naturalist school, which represent two separate poles. The intelligent design school claims that the process of existence of living things depends on very special and sensitive variables, and that this sensitive process is too complex to be explained by naturalistic reasons, and that the theory (...) of evolution was designed only by a supernatural, transcendent power. The naturalist school, on the other hand, claims that the evolution process can only be explained by natural causes and therefore there will be no need for a transcendental power. Biologist Richard Dawkins, one of the most important representatives of the naturalist school, tries to show how evolution can be explained by natural causes, based on the concept of natural selection introduced with the theory of evolution. In this research, we focused on how Dawkins explained the theory of evolution from a naturalistic perspective and examined the consistency of his conclusion on the axis of the naturalistic school paradigm that Dawkins followed. By presenting Dawkins' theory of evolution from a naturalist perspective, the position of the theory of evolution, which stands out as a scientific theory, on theological and metaphysical phenomena will be understood. At the same time, the way the naturalistic paradigm deals with scientific phenomena will be explained, based on the example of the theory of evolution. (shrink)

Charles Darwin’s theory of evolution by means of natural selection finds application far outside biology, for which it was originally invented. Its consequences for science proved far-going, influencing practically every field from thermodynamics to the humanities. While acting on biological systems, the Darwinian mechanism is a source of progress and the local-scale abandonment of the universe’s general tendency towards chaos. However, observations of changes taking place in selection-exposed organisms show that evolutionary success requires some essential limitations. The application of (...) this reasoning to social evolution may appear thought-provoking: stable evolution requires not only intense selection but also a conservative approach to ideas. (shrink)

A naturalistic theory of intentionality is proposed that differs from previous evolutionary and tracking theories. Full-blown intentionality is constructed through a series of evolvable refinements. A first, minimal version of intentionality originates from a conjectured internal process that estimates an organism’s own fitness and that continually modifies the organism. This process produces the directedness of intentionality. The internal estimator can be parsed into intentional components that point to components of the process that produces fitness. It is argued that such intentional (...) components can point to mistaken or non-existing entities. Different Fregian senses of the same reference correspond to different components that have different roles in the estimator. Intentional components that point to intentional components in other organisms produce directedness towards semi-abstract entities. Finally, adding a general, population-wide means of communication enables intentional components that point to fully abstract entities. Intentionality thus naturalized has all of its expected properties: being directed; potentially making errors; possibly pointing to non-existent, abstract, or rigid entities; capable of pointing many-to-one and one-to-many; distinguishing sense and reference; having perspective and grain; and having determinate content. Several examples, such as ‘swampman’ and ‘brain-in-a-vat’, illustrate how the theory can be applied. (shrink)

In this paper we discuss the epistemological positions of evolution theories. A sharp distinction is made between the theory that species evolved from common ancestors along specified lines of descent (here called the theory of common descent), and the theories intended as causal explanations of evolution (e.g. Lamarck's and Darwin's theory). The theory of common descent permits a large number of predictions of new results that would be improbable without evolution. For instance, (a) phylogenetic trees have been (...) validated now; (b) the observed order in fossils of new species discovered since Darwin's time could be predicted from the theory of common descent; (c) owing to the theory of common descent, the degrees of similarity and difference in newly discovered properties of more or less related species could be predicted. Such observations can be regarded as attempts to falsify the theory of common descent. We conclude that the theory of common descent is an easily-falsifiable & often-tested & still-not-falsified theory, which is the strongest predicate a theory in an empirical science can obtain. Theories intended as causal explanations of evolution can be falsified essentially, and Lamarck's theory has been falsified actually. Several elements of Darwin's theory have been modified or falsified: new versions of a theory of evolution by natural selection are now the leading scientific theories on evolution. We have argued that the theory of common descent and Darwinism are ordinary, falsifiable scientific theories. (shrink)

The target article by Denis Noble is an excellent overview of the illusions of the Modern Synthesis that still remains in textbooks despite of the recent criticism. Overcoming these illusions shows the active role of organisms in the evolutionary process and accounts for additional mechanisms such as plasticity of embryo development, epigenetic heredity, multilevel selection, Baldwin effect, and niche construction, which are components of the Extended Evolutionary Synthesis. Adding these mechanisms is certainly an important step forward, but I argue that (...) it is not sufficient for building a new theory of evolution. What is missing is a clear understanding of such notions as agency, autonomy, semiosis, interpretation, and goal-directedness, which so far belong to the humanities and have not been applied seriously in science. Organisms are autonomous and goal-directed semiotic agents capable of interpreting hereditary signs and making meaningful models of their environment. Evolutionary biology needs a semiotic vocabulary to talk about higher level functions in organisms, where specific molecules and mechanisms are only means for integrating functions over the life cycle and adapting to the environment without compromising organism integrity and identity. Such a vocabulary is being developed in biosemiotics; thus, I expect the emergence of a biosemiotic theory of evolution. (shrink)

My aims in this essay are two. First (§§1-4), I want to get clear on the very idea of a theory of the history of philosophy, the idea of an overarching account of the evolution of philosophical reflection since the inception of written philosophy. And secondly (§§5-8), I want to actually sketch such a global theory of the history of philosophy, which I call the two-streams theory.

I argue that uniquely human forms of ‘Theory of Mind’ are a product of cultural evolution. Specifically, propositional attitude psychology is a linguistically constructed folk model of the human mind, invented by our ancestors for a range of tasks and refined over successive generations of users. The construction of these folk models gave humans new tools for thinking and reasoning about mental states—and so imbued us with abilities not shared by non-linguistic species. I also argue that uniquely human forms (...) of ToM are not required for language development, such that an account of the cultural origins of ToM does not jeopardise the explanation of language development. Finally, I sketch a historical model of the cultural evolution of mental state talk. (shrink)

The book illustrates how Darwin's theory has evolved, about the development of the biological world before Darwin, and great changes that took place with the incorporation of statistics, and after Darwin's death of genetics and mathematics. The formation of 'Modern Synthesis', protein electrophoresis, Discovery of DNA opened new avenues for the study of evolution.

Charles Darwin’s theory of evolution through natural selection, explaining geographical distributions and the fossil record, is rightly regarded as one of the greatest scientific theories of all time, taking its place alongside Isaac Newton’s theory of gravitational attraction, explaining the Copernican heliocentric world picture. There is, however, a tendency to think that Darwin’s work is finished. It belongs to Victorian history rather than as something that has crucial social relevance today. This essay shows how mistaken it is to make (...) this assumption. Through a series of case histories―foreigners, class, sexual orientation, and women―Darwinism is shown to be as vibrant and important today as it was when Darwin was young. It is an essential tool for analyzing and solving some of the biggest and most pressing social issues facing us in the twenty-first century. (shrink)

Ever since proposals for generalizing the theory of natural evolution have been put forward, the aims and ambitions of both proponents and critics have differed widely. Some consider such proposals as merely metaphors, some as analogies, some aim at a real generalization and unification, and some have even proposed to work out full reductions. In this paper it is argued that these different forms of generalizing the theory of evolution can be systematically re-framed as different approaches for transferring (...) justification from the natural to the cultural realm, and that their differences are basically a matter of degree. With the help of such a classification it should become clearer what to expect, but also what not to expect from the different approaches. (shrink)

This book puts multilevel selection theory into a much needed historical perspective. This is achieved by discussing multilevel selection in the first half of the twentieth century, the reasons for the energetic rejection of Wynne-Edwards’ group selectionist stance in the 1960s, Elisabeth Lloyd’s contribution to the units of selection debate, Price’s hierarchical equation and its possible interpretations and, finally, species selection in macroevolutionary contexts. Another idea also seems to emerge from these studies; namely, that perhaps a more sure-footed position for (...) multilevel selection theory would be acquired if we were to show a renewed interest in 'old group selection', i.e. in scenarios in which the differential reproduction of the groups themselves affects the frequencies of either individual-level or group-level traits. This book will be of interest to philosophers and historians of biology, as well as to theoretically inclined biologists who have an interest in multilevel selection theory. (shrink)

The species-specific organizational property of speech is a continual mouth open-close alternation, the two phases of which are subject to continual articulatory modulation. The cycle constitutes the syllable, and the open and closed phases are segments framescontent displays that are prominent in many nonhuman primates. The new role of Broca's area and its surround in human vocal communication may have derived from its evolutionary history as the main cortical center for the control of ingestive processes. The frame and content components (...) of speech may have subsequently evolved separate realizations within two general purpose primate motor control systems: (1) a motivation-related medial system, including anterior cingulate cortex and the supplementary motor area, for self-generated behavior, formerly responsible for ancestral vocalization control and now also responsible for frames, and (2) a lateral system, including Broca's area and surround, and Wernicke's area, specialized for response to external input (and therefore the emergent vocal learning capacity) and more responsible for content. (shrink)

This paper supplements an earlier one (Wassermann 1978b). Its views aim to reinforce those of Lewontin and other prominent evolutionists, but differ significantly from the opinions of some philosophers of science, notably Popper (1957) and Olding (1978). A basic distinction is made between 'laws' and 'theories of mechanisms'. The 'Theory of Evolution' is not characterized by laws, but is viewed here as a hypertheory which explains classifiable evolutionary phenomena in terms of subordinate classifiable theories of 'evolution-specific mechanisms' (ESMs), (...) each of which could apply to a host of species. Adaptations could result from ESMs that are rooted in molecular complementarities. The status of optimization theories that aim to predict best adapted states of organisms or populations is also discussed. (shrink)

Numerous authors have examined the influence of other thinkers on Darwin’s formulation of some of the key concepts of the theory of evolution. Amongst those, Alexander von Humboldt often stands out – a scholar who, following his intention to explain the interconnection of various parts of the natural system, seems to tackle the question of evolution but does not offer an explicit answer. In this article, I examine Humboldt’s thoughts on evolution and the origin of species and (...) evaluate his contribution to Darwin’s theory of evolution. First, I analyse and explicate the fundamental assumptions and goals of Humboldt theory, and compare them to Darwin. Moving forward, I highlight the similarity of their methods, and argue that Humboldt and Darwin conduct similar investigation of fossil record and geographical distribution of species. Finally, I show that Humboldt acknowledges essential elements of Darwin’s theory of evolution: evidence given by fossil records, struggle for survival and relation between natural environment and living organisms. Humboldt, however, concludes we cannot know the evolution of species. I explain this stance, and contend that theories of Humboldt and Darwin turn out to be more similar than they seem, yet their different conclusions regarding the evolution of species stem from different initial assumptions underlying their respective frameworks. (shrink)

At the heart of evolutionary theory lays the notion of replication. Unfortunately, this notion is far less exact than the weight of its importance. In this paper, it is argued that replication always involves coding. Furthermore, when a theory of evolution is built on replication based on coding, a unifying and coherent picture arises that sheds new light on some of the controversies and open questions in contemporary biology, such as what are the roles of phylogeny and ontogeny in (...) evolution, and how to characterize and explain the increase of biological complexity. (shrink)

This article addresses the issue of human imagination from the perspective of 'niche construction' in the wider discussion about 'what makes us human' and what it means to be a 'self', specifically for the Christian faith and for theology. In the article, a brief review of human origins and human evolution demonstrates the path and substantive impact of changes in behaviour, life histories and bodies in our human ancestors and us as humans ourselves. In the interactive process of niche (...) construction, potentially changeable natural environments were, and are, acting continuously on variation in the gene pools of populations, and in this way gene pools were modified over generations. It is argued that a distinctively human imagination is part of the explanation for human evolutionary success and can be seen as one of the structurally significant aspects of the transition from earlier members of the genus Homo to ourselves as we are today. There is thus a naturalness to human imagination, even to religious imagination, that facilitates engagement with the world that is truly distinct. This provides fruitful addition to the toolkit of inquiry for both evolutionary scientists and interdisciplinary theologians interested in reconstructing the long, winding historical path to humanity. (shrink)

The Frame/Content theory deals with how and why the first language evolved the present-day speech mode of programming syllable “Frame” structures with segmental “Content” elements. The first words are considered, for biomechanical reasons, to have had the simple syllable frame structures of pre-speech babbling, and were perhaps parental terms, generated within the parent–infant dyad. Although all gestural origins theories have iconicity as a plausible alternative hypothesis for the origin of the meaning-signal link for words, they all share the problems of (...) how and why a fully fledged sign language, necessarily involving a structured phonology, changed to a spoken language. (shrink)

Drawing on his investigation of over one hundred mid-Victorian British newspapers and periodicals, Alvar Ellegård describes and analyzes the impact of Darwin's theory of evolution during the first dozen years after the publication of the Origin of Species . Although Darwin's book caused an immediate stir in literary and scientific periodicals, the popular press largely ignored it. Only after the work's implications for theology and the nature of man became evident did general publications feel compelled to react; each social (...) group responded according to his own political and religious prejudices. Ellegård charts the impact of this revolution in science, maintaining that although the idea of evolution was generally accepted, Darwin's primary contribution, the theory of natural selection, was either ignored or rejected among the public. (shrink)

Since the emergence of neo-evolutionism in the 1960s, various critiques of the theory of social or socio-cultural evolution have been forwarded, including notably those of Immanuel Wallerstein, Alain Touraine and Anthony Giddens who decisively reject the idea of evolution. Within this context, Jürgen Habermas's theory of socio-cultural evolution has also become a specific object of critique, the best known in the English-speaking world being, perhaps, Michael Schmid's critique. While the latter is ultimately based on neo-Darwinistic assumptions which (...) allow a non-Marxist alternative to be pitted against Habermas's position, a significant immanent critique has been taking shape during the late 1970s and 1980s the contours of which are at present starting to become visible, despite the fact that as yet it is nowhere developed and presented in a systematic manner. I have in mind here the work of younger authors belonging to the circle around Habermas, such as Johann Arnason, Axel Honneth, Hans Joas, Günter Frankenberg, Ulrich Rödel, and Klaus Eder, who have not only developed a critique of Habermas but are also engaged in developing an alternative to him within the framework of critical theory. (shrink)

Kary (1990) defends the view that evolution by natural selection can be adequately explained in terms of a theory incorporating only a single level of selection. Here I point out some of the inherent inadequacies of such a theory.

The paper characterizes Darwin's theory, providing a synthesis of recent historical investigations in this area. Darwin's reading of Malthus led him to appreciate the importance of population pressures, and subsequently of natural selection, with the help of the wedge metaphor. But, in itself, natural selection did not furnish an adequate account of the origin of species, for which a principle of divergence was needed. Initially, Darwin attributed this to geographical isolation, but later, following his work on barnacles which underscored the (...) significance of variation, and arising from his work on botanical arithmetic, he supposed that diversity allowed more places to be occupied in a given region. So isolation was not regarded as essential. Large regions with intense competition, and with ample variation spread by blending, would facilitate speciation. The notion of place was different from niche, and it is questioned whether Darwin's views on ecology were as modern as is commonly supposed. Two notions of struggle are found in Darwin's theory; and three notions of variation. Criticisms of his theory led him to emphasize the importance of variation over a range of forms. Hence the theory was populational rather than typological. The theory required a Lamarckian notion of inheritable changes initiated by the environment as a source of variation. Also, Darwin deployed a use/habit theory; and the notion of sexual selection. Selection normally acted at the level of the individual, though kin selection was possible. Group selection was hinted at for man. Darwin's thinking (and also the exposition of his theory) was generally guided by the domestic-organism analogy, which satisfied his methodological requirement of a vera causa principle. (shrink)

Cultural historian Iain McCalman tells the stories of Charles Darwin and his most vocal supporters and colleagues: Joseph Hooker, Thomas Huxley, and Alfred Wallace. Beginning with the somber morning of April 26, 1882--the day of Darwin's funeral--Darwin's Armada steps back in time and recounts the lives and scientific discoveries of each of these explorers. The four amateur naturalists voyaged separately from Britain to the southern hemisphere in search of adventure and scientific fame. From Darwin's inaugural trip on the Beagle in (...) 1835 through Wallace's exploits in the Amazon and Malaysia in the 1840s and 1850s, each man independently made discoveries that led him to embrace Darwin's groundbreaking theory of evolution. This book reveals the untold story of Darwin's greatest supporters who, during his life, campaigned passionately in the war of ideas over evolution and who lived on to extend and advance the scope of his work.--From publisher description. (shrink)

language to explain, and I want to show how this depends on what you think language is. So, what is language? Everybody recognizes that language is partly culturally dependent: there is a huge variety of disparate languages in the world, passed down through cultural transmission. If that’s all there is to language, a theory of the evolution of language has nothing at all to explain. We need only explain the cultural evolution of languages: English, Dutch, Mandarin, Hausa, etc. (...) are products of cultural history. However, most readers of the present volume probably subscribe to the contemporary scientific view of language, which goes beneath the cultural differences among languages. It focuses on individual language users and asks. (shrink)